- Ph.D. Marine Science. University College of North Wales, Bangor, Wales
- B.S. Biological Science. University of East Anglia, Norwich, U.K.
Physiological ecology of marine molluscs. Larval dispersal and settlement processes in estuarine and shallow water systems. Fisheries and aquaculture of marine molluscs. Invasion biology and ballast water technologies.
2003-2012, Director for Research and Advisory Services, Virginia Institute of Marine Sciences
1987-present, Professor, College of William and Mary
1985-1987, Associate Professor, College of William and Mary
1981-1984, Associate Scientist, Woods Hole Oceanographic Institution (WHOI)
1977-1981, Assistant Scientist, WHOI
1975-1977, Post-Doctoral Fellow, WHOI
- Population models of the Chesapeake oyster resources.
- Physiological and ecological studies of bivalve mollusc larvae in the field and laboratory.
- Dispersal processes in estuarine systems.
- Resource stock assessment by fishery dependent and fishery independent methods.
- Population assessment, life history and fishery management of offshore clam resources.
- Ecological restoration of oyster reef communities.
- Biology and ecology of veined rapa whelks in the Chesapeake Bay.
- Population demographics, growth rates, and age structure for Chesapeake hard clam resources.
Fellowships and Grants
7/1/2014-6/30/2015. Participation of a high school teacher in the SCeMFiS project entitled “Ocean quahogs (Arctica islandica) recruitment and life history dynamics.” - $10,000. National Science Foundation. Salary, technical and travel support to teacher participant only.
4/1/2013 – 3/31/2018. Industry/University Cooperative Research Center: Science Center for Marine Fisheries (SCeMFiS). Eric N. Powell (University of Southern Mississippi) and Roger Mann (VIMS) VIMS portion - $300,000. National Science Foundation plus $1,500,000 industry matching. Technical and travel support only from NSF portion. Variable salary from industry portion.
10/1/2014-12/31/2014. Stock assessment of Virginia Oysters. Virginia Marine Resources Commission. Field personnel support only – reimbursable to $10,000.
7/1/2012-12/30/2014. Plumeri Award for Faculty Excellence. College of William and Mary. $10,000.
7/1/2013-9/30/2014. Shell budgets in the Virginia Chesapeake Bay: Quantitative estimates to support long term restoration and resource stabilization. Virginia Oyster Heritage Foundation. Roger Mann (VIMS) and James Wesson (VMRC) $30,000 (student and technical support).
10/1/2009-9/30/2012 (no cost extension through 9/30/2014). Collaborative Research: Climate Change and Responses in a Coupled Marine System. NSF. $613,707. Lead PI McCay (Rutgers) VIMS award $99,999. Mann 1.0 mo/yr
7/1/2011-6/30/2013 (no cost extension through 6/30/2014). A bay-wide approach to oyster stock assessment, estimates of vital rates and disease status. NOAA-NCBO. $ 336,193. Mann (lead PI) 1.0 mo/yr., with Ryan Carnegie (VIMS), James Wesson (VMRC), Mike Naylor and C. Dungan (MD DNR), Kennedy Paynter Jr. (UMD) and Howard Townsend (NOAA).
2/1/2012 – 1/31/2013. Planning Grant: Industry/University Cooperative Research Center for Mid-Atlantic Center for Fisheries Science (MaCFiS). Eric N. Powell (Rutgers) and Roger Mann (VIMS) $30,000. National Science Foundation. Technical and travel support only.
1/1/2012-12/31/2013. Climate Change and the Fisheries Food Web in the Chesapeake Bay and coastal Atlantic Ocean. Robert Latour, Mary Fabrizio and Roger Mann. (VEE post doctoral support)
5/23/2011-12/21/2012. Shell substrate conditions and predator exclusion in oyster restoration. Virginia Oyster Heritage Foundation. Roger Mann (VIMS) and James Wesson (VMRC) $25,000 (technical support).
7/1/2011-6/30/2013 (no cost extension through 6/2014 to accommodate field work schedule). Oyster planting protocols to deter losses to cownose ray predation. NOAA-NCBO. $156,297. Mann (Lead PI) 1.0 mo/yr with James Wesson (VMRC), A.J. Erskine (Cowart Seafood), Tommy Leggett and Bill Goldsborough (CBF).
11/1/2010-12/31/2013 – (final report in preparation, due 3/31/2014) Shell budgets as a tool in oyster restoration and fishery management – application in Louisiana Primary State Seed Grounds. $150,000. National Fish & Wildlife Foundation. Mann (lead PI) 0.5 mo/yr.
- Hofmann, E.E., E.N. Powell, J.M. Klinck, D.M. Munroe, R. Mann, D.B. Haidvogel, D.A. NarvÁEz, X. Zhang, K.M. Kuykendall. (2018). An overview of factors affecting distribution of the Atlantic surfclam (Spisula solidissima), a continental shelf biomass dominant, during a period of climate change. J. Shellfish Res. 37:821-831, http://www.bioone.org/doi/10.2983/035.037.0412
- Pace, S. M., Powell, E. N., Mann, R., Long, M.C., and J.M. Klinck. (2017). Development of an Age-Frequency Distribution for Ocean Quahogs (Arctica islandica) on Georges Bank. J. Shellfish Res. 36: 41-53.
- Powell, E.N., Mann, R., Ashton-Alcox, K.A., Kuykendall, K.M., and M.C. Long. (2017). “Can we estimate molluscan abundance and biomass on the continental shelf?” Estuarine, Coastal and Shelf Science. 198: 213-224
- Powell, E.N., Klinck, J.M., Hofmann, E.E., Moreno, P., Kuykendall, K., Munroe, D.M., and R. Mann (2016). Captains' response to a declining stock as anticipated in the surfclam (Spisula solidissima) fishery on the U.S. Mid-Atlantic coast by model evaluation, Ocean & Coastal Management, 134, pp. 52-68. https://doi.org/10.1016/j.ocecoaman.2016.09.012
Munroe, D.; D. Narvaez; D. Hennen; E. Hofmann; L. Jacobsen; R. Mann, E. Hofmann, E.N. Powell, J. Klinck (2016). The Roles of Fishing and Bottom Water Temperature as Drivers of Change in Maximum Shell Length in Atlantic Surfclams (Spisula solidissima) Estuarine Coastal and Shelf Sciences. 170: 112–122
Powell, E.N., R. Mann, K. A, Ashton-Alcox, Y. Kim and D. Bushek (2015). The Allometry of Oysters: Spatial and Temporal Variation in the Length-Biomass Relationships for Crassostrea virginica. J. Mar. Biol. Ass. UK
- Powell, E. N., J. M. Klinck, D. M. Munroe, E. E. Hofmann, P. Moreno, R. Mann. (2015). The Value of Captains’ Behavioral Choices in the Success of the Surfclam (Spisula solidissima) Fishery on the U.S. Mid-Atlantic Coast: a Model Evaluation.
- Zhang, X., D. Haidvogel, D. M. Munroe, E. N. Powell, J. M. Klinck, R. Mann, F. Castruccio. (2015). Modeling Atlantic surfclams, Spisula solidissima, larval connectivity within the Middle Atlantic Bight: Model development, larval dispersal and metapopulation connectivity. Estuarine, Coastal and Shelf Science.
- Narváez. D. A., E. E. Hofmann, D. M. Munroe, J. M. Klinck, E. N. Powell, R. Mann, E. Curchitser. (2015). Long-term dynamics in Atlantic surfclam (Spisula Solidissima) populations: The role of bottom water temperature. J. Mar. Sci.
- Kelsey M. Kuykendall, K.M., P. Moreno, E. N. Powell, T. M. Soniat, S. Colley, R. Mann, D. M. Munroe. (2015). The exposed surface area to volume ratio: Is shell more efficient than limestone in promoting oyster recruitment? J. Shellfish Research.
- Walles, B., R. Mann, T. Ysebaert, K. Troost, P.M. J. Herman and A. Smaal (2015). Demography of the ecosystem engineer Crassostrea gigas, related to vertical reef accretion and reef persistence. Estuarine, Coastal and Shelf Science. 154: 224-233. http://dx.doi.org/1011016/j/eccs.2015.01.006.
- Soniat, T. N., N. Cooper, E. N. Powell, J. M. Klinck, M. Abdelguerfi, S. Tu, R. Mann and P. Banks. (2014). Estimating sustainable harvest of Eastern Oysters, Crassostrea virginica. J. Shellfish Research 33: 381-394. Doi:10.2983/035.033.0207.
- Mann, R., M. Southworth, R. Carnegie and R. Crockett. (2014). Temporal variation in fecundity and spawning in the eastern oyster, Crassostrea virginica, in the Piankatank River, Virginia. J. Shellfish Research 33(1): 167-176.
- Mann, R., D.M. Munroe, E.N. Powell, E.E. Hofmann and J.M. Klinck (2013). Bivalve molluscs: barometers of climate change in Arctic marine systems. In Responses of Arctic Marine Ecosystems to Climate Change. In: F.J. Mueter, D.M.S. Dickson, H.P. Huntington, J.R. Irvine, E.A. Logerwell, S.A. MacLean, L.T. Quakenbush, and C. Rosa, Alaska Sea Grant, University of Alaska Fairbanks, doi:10.4027/ramecc.2013.04.
- Harding, J. M., M. A. Unger, R. Mann, E. A. Jestel, and C. Kilduff. (2013). Rapana venosa as an indicator species for butyltin exposure over decadal and seasonal scales. Marine Biology. DOI 10.1007/s00227-013-2292-7
- Mann, R. (2013). Restoring nature’s coastal architects: a reality check. Ecological Restoration. 31(3) 323-324.
- Mann, R and M. W. Luckenbach (2013). Sperm swimming speeds in the oyster Crassostrea virginica (Gmelin 1791). J. Shellfish Research. 32(1): 1-4
- Waldbusser, G.G., E. N. Powell, and R. Mann (2013). Ecosystem effects of shell aggregations and cycling in coastal waters: An example of Chesapeake Bay oyster reefs. Ecology 94(4): 895-903
- Munroe, D.M., E.N. Powell, R. Mann, J. M. Klinck, and E. E. Hofmann. (2013). Underestimation of primary productivity on continental shelves: evidence from maximum size of extant surfclam (Spisula solidissima) populations. Fisheries Oceanography. 22: 220–233. doi: 10.1111/fog.12016
- Harding, J.M., E.N. Powell, R. Mann and M.J. Southworth. (2013). Variations in Eastern oyster (Crassostrea virginica) sex ratios from three Virginia estuaries related to growth and mortality. Jour. Mar. Biol. Assoc. U.K. 93(2): 519-531.
- Harding, J. M., M.J. Southworth, Mann, R., and J. Wesson (2012). Comparison of Crassostrea virginica Gmelin (Eastern Oyster) recruitment on constructed reefs and adjacent natural Oyster bars over decadal time scales. Northeastern Naturalist. 19(4): 627-646. DOI: http://dx.doi.org/10.1656/045.019.0407
- Harding, J.M., W. M. Walton, C. M. Trappani, M. G. Frick and R. Mann. (2011). Sea turtles as potential dispersal vectors for non-indigenous species; The Veined Rapa Whelk as an Epibiont of Loggerhead Sea Turtles. Southeastern Naturalist. 10(2): 233-244
- Kennedy, V.S., D. L. Breitburg, M. C. Christman, M. W. Luckenbach, K. Paynter, J. Kramer, K. G. Sellner, J. Dew-Baxter, C. Keller, and R. Mann. (2011). Lessons Learned from Efforts to Restore Oyster Populations in Maryland and Virginia, 1990-2007. J. Shellfish Research. 30(3):719-732
- Harding, J. M., Mann, R., M. Southworth, and J. Wesson. 2010. Management of the Piankatank River, Virginia, in support of oyster (Crassostrea virginica, Gmelin 1791) fishery repletion. J. Shellfish Research 29(4): 1-22
- Southworth, M., J. M. Harding, R. Mann and J. Wesson. 2010. Oyster (Crassostrea virginica Gmelin 1791) population dynamics on public reefs in the Great Wicomico River, Virginia, USA. J. Shellfish Research 29(2): 271-290.
- Harding, J. M., H. J. Spero, R. Mann, G. Herbert and J. Sliko. 2010. Reconstructing early 17th century estuarine drought conditions from Jamestown oysters. Proc. National Academy Sciences. 107(22) on line. www.pnas.org/cgi/doi/10.1073/pnas.1001052107
- Harding, J.M. and R. Mann. 2010. Distribution, size, and sex ratio of mature blue crabs Callinectes sapidus Rathbun 1896, from a Chesapeake Bay tributary in relation to habitat type and environmental factors. Bull. Mar Sci. 86(1): 75-91.
- Mann, R., J. M. Harding. and M. Southworth. 2009. Reconstructing pre-colonial oyster demographics in the Chesapeake Bay, USA. Estuarine, Coastal and Shelf Science 85: 217-222
- Harding, J. M., R. Mann, P. Moeller and M. S. Hsia. (2009). Rapana venosa mortality in relation to an Alexandrium monilatum bloom in the York River, USA. J. Shellfish Research. 28(2):
- Mann, R., M. Southworth, J.M. Harding and J. Wesson. (2009). Population studies of the native oyster Crassostrea virginica (Gmelin) in the James River, Virginia, USA. J. Shellfish Research. 28(2): 1-30
- Harding. J. M., R. Mann and M. Southworth. 2008. Shell length at age relationships in James River oysters (Crassostrea virginica) collected four centuries apart. J. Shellfish Research. 27(5): 1109-1115
- Harding, J. M., Gera, S. M., and R. Mann. 2008. Radula morphology in veined rapa whelks (Rapana venosa, Valenciennes 1846, Muriciade). The Nautilus 122(4):217-227
- Harding, J. M., R. Mann and C. Ware-Kilduff. 2008. Influence of environmental factors and female size on reproductive effort in a temperate marine gastropod Rapana venosa (Muricidae: Valenciennes 1846). Marine Biology 155(6): 571-581
- Harding, J. M., S. E. King. E. N. Powell, and R. Mann. (2008). Decadal trends in age structure and recruitment patterns of ocean quahogs Arctica islandica from the Mid-Atlantic Bight in relation to water temperature. J. Shellfish Research 27(4): 667-690
- Varnell, L., D.M. Bilkovic, J.Olney, D.Evans, H.Wang, C.Hershner, R. Mann. 2008. Estuarine surface water allocation: A case study on the interactive role of science in support of management. Environmental Science and Policy 11(7): 602-612.
- Harding, J.M., Mann, R, Kingsley-Smith, P and Savini, D. 2007. Comparison of predation signatures left by Atlantic oyster drills (Urosalpinx cinerea Say; Muricidae) and veined rapa whelks (Rapana venosa Valenciennes, Muricidae) in bivalve prey. Journal of Experimental Marine Biology and Ecology. 352: 1-11
- Walter, J. F. III, M. C. Christman, J. M. Hoenig, and R. Mann. 2007. Combining data from multiple years or areas to improve variogram estimation. Environmetrics 18: 1-16
- Mann, R. and E. N. Powell 2007. Why oyster restoration goals in the Chesapeake Bay are not and probably cannot be achieved. J. Shellfish Research. 26(4): 905-917.
- Harding, J.M, Mann, R. and Ware-Kilduff, C. 2007. The relationship between fecundity and size in a large marine gastropod Rapana venosa (Muricidae). Journal of Shellfish Research. 26(1):33-42.
- Olney, J.E., D. M. Bilkovic, C. H. Hershner, L. M. Varnell, H. Wang and R. Mann. 2006. Six fish and 600,000 Thirsty Folks – A Fishing Moratorium on American Shad Thwarts a Controversial Municipal Reservoir Project in Virginia USA. American Fisheries Society Symposium, 2006: 587-598.
- Mann, R, Harding, JM, and E. Westcott. 2006. Occurrence of imposex and seasonal patterns of gametogenesis in the invading veined rapa whelk Rapana venosa (Valenciennes, 1846) from Chesapeake Bay, USA. Marine Ecology Progress Series. 310: 129-138.
- Mann, R. 2006. The Local Costs To Ecological Services Associated With High Seas Global Transport. Pp 25-38 In: The Ecology of Transportation: Managing Mobility for the Environment J. Davenport and J. Davenport (eds). Kluwer Press.
- Harding, J. M. and R. Mann. 2006. Age and growth of Crassostrea ariakensis and Crassostrea gigas from Laizhou Bay, China. J. Shellfish Res. 25(1): 73-82.
- Harding, J. M. and R. Mann. 2005. Veined rapa whelk (Rapana venosa) range extensions in the Virginia waters of the Chesapeake Bay, USA. J. Shellfish Res. 381-386.
- Powell, E.N. and R. Mann. 2005. Evidence of recent recruitment in the ocean quahog Arctica islandica in the Mid-Atlantic Bight. J. Shellfish Res. 24(2): 517-530
- Mann, R., J. M. Harding, M. J. Southworth, J. A. Wesson. (2005) Hard clam Mercenaria mercenaria abundance and habitat use in Chesapeake Bay. J. Shellfish Res. 24(2): 509-516
- Mann, R., M. Southworth, J.M. Harding, and J. Wesson. 2004. A comparison of dredge and patent tongs for estimation of oyster populations. Journal of Shellfish Research. 23(2): 387-390.
- Southworth, M. and R. Mann. 2004. Decadal scale changes in seasonal patterns of oyster recruitment in the Virginia sub estuaries of the Chesapeake Bay. Journal of Shellfish Research. 23(2): 391-402.
- Mann, R. and D. Evans. 2004. Site selection for oyster habitat rehabilitation in the Virginia portion of the Chesapeake Bay: A commentary. Journal of Shellfish Research. 23(1): 41-49.
- Harding, J. M. and R. Mann. 2003. Influence of habitat on diet and distribution of striped bass (Morone saxatilis) in a temperate estuary. Bulletin of Marine Science. 72(3): 841-851.
- Mann, R. and J. M. Harding. 2003. Salinity tolerance of larval Rapana venosa: Implications for dispersal and establishment of an invading predatory gastropod on the North American Atlantic coast. Biological Bulletin. 204: 96-103.
- Baker, P.K. and R. Mann. 2003. Late stage bivalve larvae in a well-mixed estuary are not inert particles. Estuaries. 26(4A): 837-845.
- Bartol, I.K., R. Mann, and M. Vecchione. 2001. Distribution of the euryhaline squid Lolliguncula brevis in the Chesapeake Bay: Effects of selected abiotic factors. Marine Ecology Progress Series. In press.
- Harding, J.M. and R. Mann. 2001. Oyster reefs as fish habitat: Opportunistic use of restored reefs by transient fishes. Journal of Shellfish Research. 20(3): 951-959.
- Bartol, I.K, R. Mann, and M.R. Patterson. 2001. Aerobic respiratory costs of swimming in the negatively buoyant brief squid Lolliguncula brevis. Journal of Experimental Biology. 204:3639-3653.
- Mann, R. 2000. Restoring the oyster reef communities in Chesapeake Bay: a commentary. Journal of Shellfish Res. 19(1): 335-339.
- Mann, R. and J. M. Harding. 2000. Invasion of the North American Atlantic coast by a large predatory Asian mollusc. Biological Invasions. 2:7-22.
- Harding, J.M. and R. Mann. 1999. Observations on the biology of the veined rapa whelk, Rapana venosa (Valenciennes, 1846) in the Chesapeake Bay. Journal of Shellfish Research. 18(1): 9-17.
Honors, Prizes and Awards
- At-large member of the Mid-Atlantic Fisheries Management Council representing Virginia. Appointed by the Secretary of Commerce, First Term 2016-2019
- Governor's appointment to the Commonwealth's Aquaculture Advisory Board, Second Term 2013-2016
- Plumeri Faculty Award, William and Mary 2012
- Chesapeake Bay Commission Tribute to Excellence, 1994, for “...official recognition of significant contributions to the management and protection of the Chesapeake Bay.”
- School of Marine Science Outstanding Teacher Award, 2001
- Kirby Lang Visiting Professor in Marine Science, University College of North Wales, Bangor, Marine Science Laboratories, Menai Bridge, Wales, 2003
- Honored Life Member, National Shellfisheries Association, 2008
- Editor, Journal of Shellfish Research from 1982 -1986. Editorial board (current): Aquaculture (Elsevier), Journal of Shellfish Research
1. David Rudders Ph.D. 2010. (I mentored David during the final year of his work including the writing based on years of work after his major professor William DuPaul retired in 2009). Incorporating industry based dredge surveys into the assessment of the sea scallop, Placopecten magellanicus
2. Stefanie Gera. 2010. M.S. Egg Capsule Hatching Success in Rapana venosa and Urosalpinx cinerea in Relation to Temperature and Salinity.
3. Emily Chandler, M.S. 2007. Genetically monomorphic invasive populations of the rapa whelk, Rapana venosa.
4. Adriana Picariello 2006. M.S. The effects of climate change on the population biology of the surf clam, Spisula solidissima, in the Middle Atlantic Bight.
5. Elizabeth Darrow (co-chair with Mark Luckenbach,). 2004 M.S. Ecological energetics of the hard clam.
6. Doolittle, Daniel. (co-chair with Mark Patterson). 2003. M.S. Development of a method for the identification and quantification of biological targets detected by sidescan sonar
7. Catherine Ware, 2002. M.S. Temporal and spatial variation in reproductive output of the veined rapa whelk (Rapana venosa) in the Chesapeake Bay.
8. Arminda Gensler. (co-chair with John Graves). 2001, M.S. Genetic Investigations of Interspecific and Intraspecific Relationships within the Genus Rapana.
9. Erica Westcott, 2001. M.S. Seasonal reproductive activity of the invading gastropod Rapana venosa in the Chesapeake Bay
10. Rebecca Green, 2001. M.S. Morphological variation of three populations of the Veined Rapa Whelk, Rapana venosa, an invasive predatory gastropod species.
11. Juliana M. Harding. 2000. Ph.D. Ecological interactions between benthic oyster reef fishes and oysters.
12. Ian Bartol. 1999. (co chair with Mark Patterson) Ph.D. Swimming Physiology of the Brief Squid, Lolliguncula brevis.
13. Melissa Southworth, 1998. M.A. Oyster Reef Broodstock Enhancement in the Great Wicomico River, Virginia
14. Sandra Brooke, 1996, M.A. A comparison of natural and laboratory diets for the culture of marine invertebrate larvae: American Oyster, Queen Conch, and Milk Conch.
15. Elizabeth Keane - Shea, M.A. 1995 (co-chair with Mike Vecchione, Systematics Laboratory, National Museum, Smithsonian.) The early life histories of three families of cephalopods, and an examination of the concept of a paralarvae.
16. Ian Bartol, 1995, M.A. Crassostrea virginica on constructed intertidal oyster reefs: Effects of tidal height and substrate level on settlement, growth and mortality.
17. Patrick K. Baker, 1994, Ph.D. Quantification of settlement and recruitment processes in bivalve molluscs.
18. Shirley M. Baker, 1994, Ph.D. Oyster (Crassostrea virginica) metamorphosis - Effects of low oxygen.
19. Robert A. Blaylock, 1992. Ph.D. Distribution, abundance and behavior of the Cownose Ray, Rhinoptera bonasus, in Lower Chesapeake Bay.
20. L. L. Castell, 1991, M.A. Assessment of larval physiological condition using fluorescent, lipid specific stains.
21. G. Curtis Roegner, 1990, M.A. Recruitment and growth of juvenile Crassostrea virginica (Gmelin) in relation to tidal zonation.
22. Kevin McCarthy, 1990, M.A. The influence of swimming behavior of larval Crassostrea virginica (Gmelin).
23. Carrollyn Cox, 1988, M.A. Seasonal changes in the fecundity of oysters Crassostrea virginica (Gmelin) from four oyster reefs in the James River, Virginia.
24. Bernardita Campos, 1988, M.A. Swimming response of larvae of three mactrid bivalves to different salinity gradients.
25. David Eggleston, 1988, M.A. Predator-prey dynamics between the blue crab, Callinectes sapidus Rathbun, and juvenile oysters, Crassostrea virginica (Gmelin).
26. Ellen Pafford, 1988, M.A. Distribution and taxonomy of endolithic algae occurring in the shells of Crassostrea virginica in the lower James River, Virginia.
- Malacology, Larval Ecology, Biological Invasions