- Ph.D. Marine Science. University College of North Wales, Bangor, Wales
- B.S. Biological Science. University of East Anglia, Norwich, U.K.
Physiological ecology of marine molluscs. Larval dispersal and settlement processes in estuarine and shallow water systems. Fisheries and aquaculture of marine molluscs. Invasion biology and ballast water technologies.
2003-2012, Director for Research and Advisory Services, Virginia Institute of Marine Sciences
1987-present, Professor, College of William and Mary
1985-1987, Associate Professor, College of William and Mary
1981-1984, Associate Scientist , Woods Hole Oceanographic Institution (WHOI)
1977-1981, Assistant Scientist , WHOI
1975-1977, Post-Doctoral Fellow, WHOI
- Population models of the Chesapeake oyster resources.
- Physiological and ecological studies of bivalve mollusc larvae in the field and laboratory.
- Dispersal processes in estuarine systems.
- Resource stock assessment by fishery dependent and fishery independent methods.
- Population assessment, life history and fishery management of offshore clam resources.
- Ecological restoration of oyster reef communities.
- Biology and ecology of veined rapa whelks in the Chesapeake Bay.
- Population demographics, growth rates, and age structure for Chesapeake hard clam resources.
Fellowships and Grants
7/1/2011-6/30/2012. A bay-wide approach to oyster stock assessment, estimates of vital rates and disease status. NOAA-NCBO.
7/1/2011-6/30/2013. Oyster planting protocols to deter losses to cownose ray predation. NOAA-NCBO.
7/1/2010-12/31/2011. Collaborative research: summer flounder collections for regional examination of sex ratio and size. Rutgers University.
7/1/2010 – 6/30/2012. Virginia oyster management and restoration. NOAA-NCBO.
11/1/2010-10/31/2012. Shell budgets as a tool in oyster restoration and fishery management – application in Louisiana Primary State Seed Grounds. National Fish & Wildlife Foundation.
2/1/2009-1/31/2012. Climate Change Impacts in Virginia: Natural resource data records as tools to assess continuing trends. NOAA Roger Mann, Marcia Berman and Carl Hershner.
10/1/2009-9/30/2012. Collaborative Research: Climate Change and Responses in a Coupled Marine System. NSF. $613,707. Lead PI Bonnie McCay (Rutgers) VIMS award.
- Kennedy, V.S., D. L. Breitburg, M. C. Christman, M. W. Luckenbach, K. Paynter, J. Kramer, K. G. Sellner, J. Dew-Baxter, C. Keller, and R. Mann. (in review). Lessons Learned from Efforts to Restore Oyster Populations in Maryland and Virginia, 1990-2007. submitted to J. Shellfish Research.
- Harding, J.M., W. M. Walton, C. M. Trappani, M. G. Frick and R. Mann. (in press). Sea turtles as potential dispersal vectors for non-indigenous species. Southeastern Naturalist.
- Harding, J. M., Mann, R., M. Southworth, and J. Wesson. 2010. Management of the Piankatank River, Virginia, in support of oyster (Crassostrea virginica, Gmelin 1791) fishery repletion. J. Shellfish Research 29(4): 1-22
- Southworth, M., J. M. Harding, R. Mann and J. Wesson. 2010. Oyster (Crassostrea virginica Gmelin 1791) population dynamics on public reefs in the Great Wicomico River, Virginia, USA. J. Shellfish Research 29(2): 271-290.
- Harding, J. M., H. J. Spero, R. Mann, G. Herbert and J. Sliko. 2010. Reconstructing early 17th century estuarine drought conditions from Jamestown oysters. Proc. National Academy Sciences. 107(22) on line. www.pnas.org/cgi/doi/10.1073/pnas.1001052107
- Harding, J.M. and R. Mann. 2010. Distribution, size, and sex ratio of mature blue crabs Callinectes sapidus Rathbun 1896, from a Chesapeake Bay tributary in relation to habitat type and environmental factors. Bull. Mar Sci. 86(1): 75-91.
- Mann, R., J. M. Harding. and M. Southworth. 2009. Reconstructing pre-colonial oyster demographics in the Chesapeake Bay, USA. Estuarine, Coastal and Shelf Science 85: 217-222
- Harding, J. M., R. Mann, P. Moeller and M. S. Hsia. (2009). Rapana venosa mortality in relation to an Alexandrium monilatum bloom in the York River, USA. J. Shellfish Research. 28(2):
- Mann, R., M. Southworth, J.M. Harding and J. Wesson. (2009). Population studies of the native oyster Crassostrea virginica (Gmelin) in the James River, Virginia, USA. J. Shellfish Research. 28(2): 1-30
- Harding. J. M., R. Mann and M. Southworth. 2008. Shell length at age relationships in James River oysters (Crassostrea virginica) collected four centuries apart. J. Shellfish Research. 27(5): 1109-1115
- Harding, J. M., Gera, S. M., and R. Mann. 2008. Radula morphology in veined rapa whelks (Rapana venosa, Valenciennes 1846, Muriciade). The Nautilus 122(4):217-227
- Harding, J. M., R. Mann and C. Ware-Kilduff. 2008. Influence of environmental factors and female size on reproductive effort in a temperate marine gastropod Rapana venosa (Muricidae: Valenciennes 1846). Marine Biology 155(6): 571-581
- Harding, J. M., S. E. King. E. N. Powell, and R. Mann. (2008). Decadal trends in age structure and recruitment patterns of ocean quahogs Arctica islandica from the Mid-Atlantic Bight in relation to water temperature. J. Shellfish Research 27(4): 667-690
- Varnell, L., D.M. Bilkovic, J.Olney, D.Evans, H.Wang, C.Hershner, R. Mann. 2008. Estuarine surface water allocation: A case study on the interactive role of science in support of management. Environmental Science and Policy 11(7): 602-612.
- Harding, J.M., Mann, R, Kingsley-Smith, P and Savini, D. 2007. Comparison of predation signatures left by Atlantic oyster drills (Urosalpinx cinerea Say; Muricidae) and veined rapa whelks (Rapana venosa Valenciennes, Muricidae) in bivalve prey. Journal of Experimental Marine Biology and Ecology. 352: 1-11
- Walter, J. F. III, M. C. Christman, J. M. Hoenig, and R. Mann. 2007. Combining data from multiple years or areas to improve variogram estimation. Environmetrics 18: 1-16
- Mann, R. and E. N. Powell 2007. Why oyster restoration goals in the Chesapeake Bay are not and probably cannot be achieved. J. Shellfish Research. 26(4): 905-917.
- Harding, J.M, Mann, R. and Ware-Kilduff, C. 2007. The relationship between fecundity and size in a large marine gastropod Rapana venosa (Muricidae). Journal of Shellfish Research. 26(1):33-42.
- Olney, J.E., D. M. Bilkovic, C. H. Hershner, L. M. Varnell, H. Wang and R. Mann. 2006. Six fish and 600,000 Thirsty Folks – A Fishing Moratorium on American Shad Thwarts a Controversial Municipal Reservoir Project in Virginia USA. American Fisheries Society Symposium, 2006: 587-598.
- Mann, R, Harding, JM, and E. Westcott. 2006. Occurrence of imposex and seasonal patterns of gametogenesis in the invading veined rapa whelk Rapana venosa (Valenciennes, 1846) from Chesapeake Bay, USA. Marine Ecology Progress Series. 310: 129-138.
- Mann, R. 2006. The Local Costs To Ecological Services Associated With High Seas Global Transport. Pp 25-38 In: The Ecology of Transportation: Managing Mobility for the Environment J. Davenport and J. Davenport (eds). Kluwer Press.
- Harding, J. M. and R. Mann. 2006. Age and growth of Crassostrea ariakensis and Crassostrea gigas from Laizhou Bay, China. J. Shellfish Res. 25(1): 73-82.
- Harding, J. M. and R. Mann. 2005. Veined rapa whelk (Rapana venosa) range extensions in the Virginia waters of the Chesapeake Bay, USA. J. Shellfish Res. 381-386.
- Powell, E.N. and R. Mann. 2005. Evidence of recent recruitment in the ocean quahog Arctica islandica in the Mid-Atlantic Bight. J. Shellfish Res. 24(2): 517-530
- Mann, R., J. M. Harding, M. J. Southworth, J. A. Wesson. (2005) Hard clam Mercenaria mercenaria abundance and habitat use in Chesapeake Bay. J. Shellfish Res. 24(2): 509-516
- Mann, R., M. Southworth, J.M. Harding, and J. Wesson. 2004. A comparison of dredge and patent tongs for estimation of oyster populations. Journal of Shellfish Research. 23(2): 387-390.
- Southworth, M. and R. Mann. 2004. Decadal scale changes in seasonal patterns of oyster recruitment in the Virginia sub estuaries of the Chesapeake Bay. Journal of Shellfish Research. 23(2): 391-402.
- Mann, R. and D. Evans. 2004. Site selection for oyster habitat rehabilitation in the Virginia portion of the Chesapeake Bay: A commentary. Journal of Shellfish Research. 23(1): 41-49.
- Harding, J. M. and R. Mann. 2003. Influence of habitat on diet and distribution of striped bass (Morone saxatilis) in a temperate estuary. Bulletin of Marine Science. 72(3): 841-851.
- Mann, R. and J. M. Harding. 2003. Salinity tolerance of larval Rapana venosa: Implications for dispersal and establishment of an invading predatory gastropod on the North American Atlantic coast. Biological Bulletin. 204: 96-103.
- Baker, P.K. and R. Mann. 2003. Late stage bivalve larvae in a well-mixed estuary are not inert particles. Estuaries. 26(4A): 837-845.
- Bartol, I.K., R. Mann, and M. Vecchione. 2001. Distribution of the euryhaline squid Lolliguncula brevis in the Chesapeake Bay: Effects of selected abiotic factors. Marine Ecology Progress Series. In press.
- Harding, J.M. and R. Mann. 2001. Oyster reefs as fish habitat: Opportunistic use of restored reefs by transient fishes. Journal of Shellfish Research. 20(3): 951-959.
- Bartol, I.K, R. Mann, and M.R. Patterson. 2001. Aerobic respiratory costs of swimming in the negatively buoyant brief squid Lolliguncula brevis. Journal of Experimental Biology. 204:3639-3653.
- Mann, R. 2000. Restoring the oyster reef communities in Chesapeake Bay: a commentary. Journal of Shellfish Res. 19(1): 335-339.
- Mann, R. and J. M. Harding. 2000. Invasion of the North American Atlantic coast by a large predatory Asian mollusc. Biological Invasions. 2:7-22.
- Harding, J.M. and R. Mann. 1999. Observations on the biology of the veined rapa whelk, Rapana venosa (Valenciennes, 1846) in the Chesapeake Bay. Journal of Shellfish Research. 18(1): 9-17.
Honors, Prizes and Awards
Chesapeake Bay Commission Tribute to Excellence, 1994, for “...official recognition of significant contributions to the management and protection of the Chesapeake Bay.”
School of Marine Science Outstanding Teacher Award, 2001
Kirby Lang Visiting Professor in Marine Science, University College of North Wales, Bangor, Marine Science Laboratories, Menai Bridge, Wales, 2003
Honored Life Member, National Shellfisheries Association, 2008
Editor, Journal of Shellfish Research from 1982 -1986.
Editorial board (current): Aquaculture (Elsevier), Journal of Shellfish Research
1. Stefanie Gera. 2010 M.S. Egg Capsule Hatching Success in Rapana venosa andUrosalpinx cinerea in Relation to Temperature and Salinity.
2. Adriana Picariello 2006. M.S. The effects of climate change on the population biology of the surf clam, Spisula solidissima, in the Middle Atlantic Bight.
3. Elizabeth Darrow (co-chair with Mark Luckenbach,). 2004 M.S. Ecological energetics of the hard clam.
4. Doolittle, Daniel. (co-chair with Mark Patterson). 2003. M.S. Development of a method for the identification and quantification of biological targets detected by sidescan sonar
5. Catherine Ware, 2002. M.S. Temporal and spatial variation in reproductive output of the veined rapa whelk (Rapana venosa) in the Chesapeake Bay.
6. Arminda Gensler. (co-chair with John Graves). 2001, M.S. Genetic Investigations of Interspecific and Intraspecific Relationships within the Genus Rapana.
7. Erica Westcott, 2001 M.S. Seasonal reproductive activity of the invading gastropod Rapana venosa in the Chesapeake Bay
8. Rebecca Green, 2001. M.S. Morphological variation of three populations of the Veined Rapa Whelk, Rapana venosa, an invasive predatory gastropod species.
9. Juliana M. Harding. 2000. Ph.D. Ecological interactions between benthic oyster reef fishes and oysters.
10. Ian Bartol. 1999. (co chair with Mark Patterson) Ph.D. Swimming Physiology of the Brief Squid, Lolliguncula brevis.
11. Melissa Southworth, 1998. M. A. Oyster Reef Broodstock Enhancement in the Great Wicomico River, Virginia
12. Sandra Brooke, 1996, M.A. A comparison of natural and laboratory diets for the culture of marine invertebrate larvae: American Oyster, Queen Conch, and Milk Conch.
13. Elizabeth Keane - Shea, M.A. 1995 (co-chair with Mike Vecchione, Systematics Laboratory, National Museum, Smithsonian.) The early life histories of three families of cephalopods, and an examination of the concept of a paralarvae.
14. Ian Bartol, 1995, M.A. Crassostrea virginica on constructed intertidal oyster reefs: Effects of tidal height and substrate level on settlement, growth and mortality.
15. Patrick K. Baker, 1994, Ph.D. Quantification of settlement and recruitment processes in bivalve molluscs.
16. Shirley M. Baker, 1994, Ph.D. Oyster (Crassostrea virginica) metamorphosis - Effects of low oxygen.
17. Robert A. Blaylock, 1992. Ph.D. Distribution, abundance and behavior of the Cownose Ray, Rhinoptera bonasus, in Lower Chesapeake Bay.
18. L. L. Castell, 1991, M.A. Assessment of larval physiological condition using fluorescent, lipid specific stains.
19. G. Curtis Roegner, 1990, M.A. Recruitment and growth of juvenile Crassostrea virginica (Gmelin) in relation to tidal zonation.
20. Kevin McCarthy, 1990, M.A. The influence of swimming behavior of larval Crassostrea virginica (Gmelin).
21. Carrollyn Cox, 1988, M.A. Seasonal changes in the fecundity of oysters Crassostrea virginica (Gmelin) from four oyster reefs in the James River, Virginia.
22. Bernardita Campos, 1988, M.A. Swimming response of larvae of three mactrid bivalves to different salinity gradients.
23. David Eggleston, 1988, M.A. Predator-prey dynamics between the blue crab, Callinectes sapidus Rathbun, and juvenile oysters, Crassostrea virginica (Gmelin).
24. Ellen Pafford, 1988, M.A. Distribution and taxonomy of endolithic algae occurring in the shells of Crassostrea virginica in the lower James River, Virginia.
- Malacology, Larval Ecology, Biological Invasions